Under Construction

Coptoclavidaegroup is extinct

David R. Maddison
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Containing group: Adephaga

Notes

This Mesozoic family consists of medium-sized to large aquatic beetles with divided compound eyes. Larvae have raptorial front legs. The subfamily Necronectinae is found in the Middle to Late Jurassic of Asia and Europe, and Early Cretaceous of northern Africa, Coptoclavinae in the Early Cretaceous of Asia, and Charonoscaphinae in the Late Jurassic of Asia. See Ponomarenko (1977, 1992) for further details.

 

Larvae

The following pictures are of larvae of the genus Coptoclava, from the Early Cretaceous of the Shandong Province of China.

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Head

Paddle-like mesothoracic and metathoracic legs

End of abdomen

References

Ponomarenko, A.G. 1977. Suborder Adephaga. Pp. 17-104 in Arnold'di, L.V., V.V. Zherikhin, L.M. Nikritin, and A.G. Ponomarenko (eds.), Mezozoiskie Zhestkokrylye. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta, Vol. 161. Nauka Publishers, Moscow.

Ponomarenko, A.G. 1992. Suborder Adephaga. Pp. 19-142 in Arnold'di, L.V., V.V. Zherikhin, L.M. Nikritin, and A.G. Ponomarenko (eds.), Mesozoic Coleoptera. Smithsonian Institution Libraries, Washington, D.C. (English translation of Ponomarenko, 1977.)

Title Illustrations
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Scientific Name Coptoclava
Location Shandong Province, China
Specimen Condition Fossil -- Period: Early Cretaceous
Life Cycle Stage larva
Image Use creative commons This media file is licensed under the Creative Commons Attribution License - Version 3.0.
Copyright © 1995 David R. Maddison
About This Page

David R. Maddison
Oregon State University

Page: Tree of Life Coptoclavidae. Authored by David R. Maddison. The TEXT of this page is licensed under the Creative Commons Attribution License - Version 3.0. Note that images and other media featured on this page are each governed by their own license, and they may or may not be available for reuse. Click on an image or a media link to access the media data window, which provides the relevant licensing information. For the general terms and conditions of ToL material reuse and redistribution, please see the Tree of Life Copyright Policies.

Citing this page:

Maddison, David R. 1995. Coptoclavidae. Version 01 January 1995 (under construction). http://tolweb.org/Coptoclavidae/8896/1995.01.01 in The Tree of Life Web Project, http://tolweb.org/

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